Advanced Poodle GeneticsVolume II: Merle & Dilution
The Science Behind Merle and Dilution Genetics
The merle genetics book written for breeders who need more than a carrier/clear label. A deep dive into the M locus, dilution genetics, and how to read a DNA panel that actually reports mosaic profiles, referenced against the current peer-reviewed literature.
- 301 pages
- The complete science of the M locus and double-merle risk
- Dilution genetics: d1, d2, and d3
- Interpreting DNA test results, including mosaic profiles
- An evidence-based look at merle’s disputed origin in Poodles

Inside Volume II
301 pages, entirely focused on merle and dilution genetics in Poodles.
The M Locus, In Full
The complete genetic picture of the merle allele: how it works, why it is reported as base-pair lengths rather than a simple carrier state, and what that means for a breeding decision.
Double-Merle Risk
The health risks that come from combining two merle alleles, covered with the seriousness the topic requires.
Dilution Genetics: d1, d2, d3
How the dilution series works and how it interacts with the rest of a poodle’s color genotype.
Reading a Real DNA Panel
How to interpret DNA test results in practice, including mosaic profiles that a simple clear/carrier readout doesn’t capture.
Merle’s Disputed Origin in Poodles
An evidence-based look at where merle in Poodles is thought to come from, referenced against the current peer-reviewed literature.

Ready to Master Poodle Genetics?
Three volumes, one continuous body of work: coat colour, merle & dilution, and health & structural genetics. Buy each on its own, or get all three together for less than the price of two.
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Xyon Marley
Author & Educator
Written for Breeders, Backed by Research
Xyon Marley is a canine genetics educator and author. He writes and publishes under the Poodle Genetics Lab imprint, an independent, breeder-neutral resource focused on the genetics that actually shape a breeding program, grounded in the current peer-reviewed literature rather than folklore.
He built Poodle Genetics Lab because the science of coat colour inheritance sat locked inside dense academic papers in journals like Animal Genetics and PLOS Genetics, out of reach of the breeders and owners who need it most. The Poodle Color Genetics series is the result: a multi-volume scientific reference that translates that research into decisions breeders can make with confidence.